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Leaf development requires highly regulated cell proliferation,differentiation and expansion patterns.As a model legume species,Medicago truncatula exhibits the simplest form of compound leaves; a single terminal leaflet is developed on the distal end of rachis and a pair of lateral leaflets is developed on both sides of petiole.M.truncatula belongs to the inverted repeat lacking clade(IRLC)of legumes.In the IRLC species,the orthologs of LEAFY function in place of KNOX1 to regulate compound leaf development,implying that the regulation mechanism of leaf development in M.