Pollen tube guidance is a common process in angiosperm that involves cell-cell interactions and signaling between the male gametophyte (pollen) and the female gametophyte.The putative female cues to attract pollen tube have been shown to be Cys-rich defensin-like peptide secreted by the synergid.We previously showed that the central cell plays an important role in guidance as mutation in CCG abolishes this process.