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Regulation of gene expression is associated with RNA interference(RNAi)at the post-transcriptional level and chromatin modification in transcriptional silencing during plant development.Dicer-like(DCL)and Argonaute(AGO)proteins,along with RNA-dependent RNA polymerase(RDR)play integral roles in these processes.Many members of the DCL,AGO,and RDR gene families have been characterized in Arabidopsis,rice,maize and tomato.