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Although studies in the past showed that embryo and endosperm developments are two parallel and well-coordinated developmental processes in seed formation, signals involved in the embryo-endosperm communication are largely unknown.We showed in this study that CLAVATA3/ESR-RELATED (CLE) 19 in Arabidopsis is expressed specifically in epidermal layers of cotyledon primordia and the lower portion of the embryo, but not in endosperm.