As one of the most important vegetables,tomato (Solanum lyco-persicum) is extensively produced and consumed worldwide and substantially contributes to human nutrition and health (The Tomato Genome Consortium,2012).Although red tomatoes are the most common,pink tomatoes are more popular in Asia,particularly in China and Japan,because of their better taste (Ballester et al.,2010;Zhu et al.,2018).Compared with red tomatoes,pink tomatoes fail to accumulate the yellow-colored flavonoid pigment,naringenin chalcone (NarCh),in their peels,resulting in a colorless peel phenotype (Adato et al.,2009;Ballester et al.,2010).Genetic studies have demonstrated that this colorless peel trait results from loss-of-function in MYB12,a master transcription factor regulating flavonoid biosynthesis (Adato et al.,2009;Ballester et al.,2010).To date,five null alleles of MYB12 have been identified and used for pink-fruited tomato breeding by the advanced backcross strategy (Fandez-Moreno et al.,2016).