Plant development in its majority occurs post-embryonically through the activity of local meristems that provide daughter cells for the development of new organs.It has long been acknowledged that the shoot apical meristem(SAM),which holds the stem cells that will form above-ground organs,is recalcitrant to infection by multiple pathogens,a crucial strategy to safeguard normal devel-opment and subsequent generations.However,the molecular mechanisms underlying SAM immunity remain largely unknown.