Meyer利用将野生二倍体棉花哈克尼西棉(G.harknessii Brandg.)的细胞质转移到四倍体种上的方法,诱导出陆地棉的细胞质雄性不育。哈克尼西棉的细胞质使栽培棉花具有完全的雄性不育性。哈克尼西棉中的恢复基因Rf 被认为是不完全显性(Weaver和Weaver)。要求遗传性的改良成分能恢复完全的雄性可育性,发现海岛棉是这种改良成份等位基因的一个来源。 Meyer uses cytoplasm of the wild diploid cotton G. harknessii Brandg. To transfer to tetraploid species to induce cytoplasmic male sterility in upland cotton. The cytoplasm of Hackney West cotton gives full male sterility to the cultivated cotton. The Rf gene in Hackney West cotton is considered incompletely dominant (Weaver and Weaver). Requiring hereditary modifications to restore complete male fertility, island cotton was found to be a source of such improved alleles.