The many roles of jasmonate in plant growth,reproduction,and defense raise interesting questions about how this lipid-derived hormone emerged during evolution.Recent studies indicate that a "higher plant" pathway involving jasmonoyl-L-isoleucine (JA-lle) as the receptor-active ligand is conserved in vascular plants (Bowman et al.,2017;Howe et al.,2018).Within this core signaling pathway,JAZ proteins link JA-Ile perception to changes in the activity of MYC transcription factors (TFs) (Figure 1) and have emerged as a conserved hub for crosstalk in hormone response pathways.Therefore,understanding how JAZ proteins work across the plant tree of life is relevant to broad areas of plant biology.