灰葡萄孢是一种重要的植物病原真菌,其寄主范围广泛,能危害世界上230多种双子叶植物,常给农业生产造成重大的经济损失[1 3]。由灰葡萄孢引起的灰霉病是目前我国温室蔬菜生产中最主要的病害之一,一般造成全年减产20%25%,严重时达到40%以上[4]。因此,研究该病菌的致病机理对该病防治具有重要意义,并且随着灰葡萄孢基因组测序的完成,灰葡萄孢已成为发育生物学、分子植物病理学研究的模式生 Botrytis cinerea is an important phytopathogenic fungus with a wide range of hosts that can endanger more than 230 dicotyledons of the world and often cause major economic losses to agricultural production [13]. Gray mold caused by Botrytis cinerea is one of the most important diseases in the production of greenhouse vegetables in our country at present, generally causing 20% ​​25% loss in the whole year and 40% in severe cases [4]. Therefore, studying the pathogenic mechanism of this bacterium is of great significance for the prevention and treatment of this disease. With the completion of the genome sequencing of Botrytis cinerea, Botrytis cinerea has become a model for developmental biology and molecular phytopathology