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The genome of Tobacco bushy top virus (TBTV) has a 4152 nt of positive, single-strand RNA, which lacks 5cap and 3poly (A) tail.Based on software prediction and homology alignment,TBTV encodes 4 proteins (5→3) : ORF1 and ORF2, which are the functional proteins involved in the replication of genome;the highly overlapped ORF3 and ORF4, which are possibly expressed from a subgenomic RNA.ORF2 was expressed as ORF1/ORF2 fused pattern by translational frameshift, which functions as RdRp.